{"id":3701,"date":"2026-09-09T22:55:52","date_gmt":"2026-09-09T22:55:52","guid":{"rendered":"https:\/\/keyapa.com\/sp\/?p=3701"},"modified":"2026-09-09T23:08:27","modified_gmt":"2026-09-09T23:08:27","slug":"defining-grasses-the-amazing-awns-of-grasses","status":"publish","type":"post","link":"https:\/\/keyapa.com\/sp\/2026\/09\/09\/defining-grasses-the-amazing-awns-of-grasses\/","title":{"rendered":"Defining Grasses: The Amazing Awns of Grasses"},"content":{"rendered":"\n<figure class=\"wp-block-image aligncenter size-full\"><img fetchpriority=\"high\" decoding=\"async\" width=\"900\" height=\"699\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0116.jpg\" alt=\"Arrhenatherum elatius in Copenhagen.\" class=\"wp-image-1097\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0116.jpg 900w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0116-300x233.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0116-768x596.jpg 768w\" sizes=\"(max-width: 900px) 100vw, 900px\" \/><figcaption class=\"wp-element-caption\"><em>Arrhenatherum elatius<\/em> in Copenhagen.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">If one were in the habit of identifying grasses, then the presence of visible and distinctive structures on the spikelets would be one way to  ease the pain of such an endeavor. Fortunately, many grasses have long projections from these reproductive structures that go a long way towards helping the observer. These so-called <strong>awns<\/strong> are long bristle-like projections on the floral bracts of grass spikelets that are structurally diverse and widespread. Although they are found in a few other plants, the Poaceae have the most diverse and abundant panoply of awns.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img decoding=\"async\" width=\"716\" height=\"667\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/grass_parts.jpg\" alt=\"Spikelets with awns (from Petersen and Kellogg. 2022).\" class=\"wp-image-3719\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/grass_parts.jpg 716w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/grass_parts-300x279.jpg 300w\" sizes=\"(max-width: 716px) 100vw, 716px\" \/><figcaption class=\"wp-element-caption\">Spikelets with awns (from Petersen and Kellogg. 2022).<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Most awns arise from the lemmas of a spikelet, usually from the mid-vein, but some originate from the glumes, and more rarely from the palea. Many are straight, others are geniculate (bent), and two other types are specific towards some taxa. The Aristida for example, have coiled awns, while the Streptochaeta have branched awns.  <\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img decoding=\"async\" width=\"900\" height=\"602\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1032.jpg\" alt=\"Avena fatua with geniculate (bent) awns rising from the lemma.\" class=\"wp-image-3715\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1032.jpg 900w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1032-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1032-768x514.jpg 768w\" sizes=\"(max-width: 900px) 100vw, 900px\" \/><figcaption class=\"wp-element-caption\"><em>Avena fatua<\/em> with geniculate (bent) awns rising from the lemma.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">The function of awns has been studied but still presents some mysteries. The most obvious is that they are involved in reproduction. In the case of grasses, the awns might aid in dispersal of the diaspore (the dispersal unit of grasses). These can be grouped into three types.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">(1) The awns aid in making sure seeds drop straight down to the ground, similar to how a parachute can stabilize a dropping package. This is important to grasses because having the base of the seeds (the callus) grounded increases germination. <\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"800\" height=\"535\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/04\/DSC_6847.jpg\" alt=\"Piptochaetium sp (Speargrass) with long straight awns at Wekiwa Springs State Park.\" class=\"wp-image-2942\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/04\/DSC_6847.jpg 800w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/04\/DSC_6847-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/04\/DSC_6847-768x514.jpg 768w\" sizes=\"(max-width: 800px) 100vw, 800px\" \/><figcaption class=\"wp-element-caption\"><em>Piptochaetium<\/em> sp (Speargrass) with long straight awns at Wekiwa Springs State Park.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">(2) The awns may also allow attachment of the diaspore to passing animals, who then carry it far from the original location. One is easily reminded of this when wading through a field of grass and emerging with many grass seeds embedded in clothing, or bemoaning the sight of a pet dog dappled with sticky particles on their fur.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"900\" height=\"602\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/09\/DSC_0966.jpg\" alt=\"Nassella tenuissima spikelets with long straight awns in Cerritos, California.\" class=\"wp-image-1682\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/09\/DSC_0966.jpg 900w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/09\/DSC_0966-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/09\/DSC_0966-768x514.jpg 768w\" sizes=\"(max-width: 900px) 100vw, 900px\" \/><figcaption class=\"wp-element-caption\"><em>Nassella tenuissima<\/em> spikelets with long straight awns in Cerritos, California.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">(3) The most dramatic way they aid in dispersal are in those seeds that have hygroscopic awns that twist  and untwist in response to changes in moisture. The movement of the awns allow the seeds to either walk across the ground (see video below).<\/p>\n\n\n\n<figure class=\"wp-block-embed is-type-video is-provider-youtube wp-block-embed-youtube wp-embed-aspect-16-9 wp-has-aspect-ratio\"><div class=\"wp-block-embed__wrapper\">\n<iframe title=\"These Seeds Can Walk! | The Green Planet | BBC Earth\" width=\"500\" height=\"281\" src=\"https:\/\/www.youtube.com\/embed\/NlUparIDfzE?feature=oembed\" frameborder=\"0\" allow=\"accelerometer; autoplay; clipboard-write; encrypted-media; gyroscope; picture-in-picture; web-share\" referrerpolicy=\"strict-origin-when-cross-origin\" allowfullscreen><\/iframe>\n<\/div><figcaption class=\"wp-element-caption\">Walking Seeds<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Just as remarkable, similar movements can enable these awned seeds to actively dig themselves deeper into the soil in order to increase their chances of germinating successfully! <\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"684\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1605-1024x684.jpg\" alt=\"Aristida beyrichiana (A. stricta) with twisted awns.\" class=\"wp-image-3707\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1605-1024x684.jpg 1024w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1605-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1605-768x513.jpg 768w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1605.jpg 1200w\" sizes=\"(max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"wp-element-caption\"><em>Aristida beyrichiana<\/em> (<em>A. stricta<\/em>) with twisted awns.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">In addition to aiding in dispersal, awns can also protect the diaspores from predatory animals. The rigid and sharp structures not only deter larger animals from ingesting them, but they also increase handling time for smaller insects, who may opt for less troublesome seeds.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"809\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1602-1024x809.jpg\" alt=\"Aristida beyrichiana (A. stricta) with coiled awns.\" class=\"wp-image-3706\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1602-1024x809.jpg 1024w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1602-300x237.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1602-768x607.jpg 768w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1602.jpg 1200w\" sizes=\"(max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"wp-element-caption\"><em>Aristida beyrichiana<\/em> (<em>A. stricta<\/em>) with coiled awns.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Another possible indirect function of the awns is to protect the diaspores from fire. This is particularly the case for hygroscopic awns that dig the seeds deep into the soil, where they are protected from the worst effects of surface fires.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"854\" height=\"668\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1632.jpg\" alt=\"Andropogon ternarius (syn.\u00a0Andropogon cabanisii)\" class=\"wp-image-3709\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1632.jpg 854w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1632-300x235.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1632-768x601.jpg 768w\" sizes=\"(max-width: 854px) 100vw, 854px\" \/><figcaption class=\"wp-element-caption\"><em>Andropogon ternarius (syn.&nbsp;Andropogon cabanisii)<\/em><\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Finally, in some grass lineages like rice (<em>Oryza<\/em> spp) and wheat (<em>Triticum<\/em> spp) the awns actively photosynthesize and supply carbon to the developing seeds, although in many grasses like those in the Andropogoneae this function is done by a nearby sterile spikelet.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"1000\" height=\"668\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1499.jpg\" alt=\"The long stiff awns of Sorghastrum secundum.\" class=\"wp-image-3711\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1499.jpg 1000w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1499-300x200.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1499-768x513.jpg 768w\" sizes=\"(max-width: 1000px) 100vw, 1000px\" \/><figcaption class=\"wp-element-caption\">The long stiff awns of <em>Sorghastrum secundum<\/em>.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Awns in the Poaceae did not arise from the common ancestor of the family, but rather seemed to have been a result of convergent evolution, with awns evolving independently at least twelve times during the history of the group.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"1000\" height=\"1013\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1351.jpg\" alt=\"The long stiff awns of Aegylops triuncialis are covered with minute barbs that may help in attaching the diaspores to passing animals. They can also physically injure large animals.\" class=\"wp-image-3712\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1351.jpg 1000w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1351-296x300.jpg 296w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_1351-768x778.jpg 768w\" sizes=\"(max-width: 1000px) 100vw, 1000px\" \/><figcaption class=\"wp-element-caption\">The long stiff awns of <em>Aegylops triuncialis<\/em> are covered with minute barbs that may help in attaching the diaspores to passing animals. They can also physically injure large animals.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">The next time you examine some grass inflorescence, take a closer look whether it has awns and see whether you can guess what their function it might have for that species.  Below are some other images of awns in different species.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"1000\" height=\"668\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_0650.jpg\" alt=\"Avena fatua spikelets in Ollantaytambo, Peru\" class=\"wp-image-3713\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_0650.jpg 1000w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_0650-300x200.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2026\/09\/DSC_0650-768x513.jpg 768w\" sizes=\"(max-width: 1000px) 100vw, 1000px\" \/><figcaption class=\"wp-element-caption\"><em><em>Avena fatua<\/em> spikelets <\/em>in Ollantaytambo, Peru<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"800\" height=\"535\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/10\/DSC_6117b.jpg\" alt=\"Oplismeus burmanni.\" class=\"wp-image-1916\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/10\/DSC_6117b.jpg 800w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/10\/DSC_6117b-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/10\/DSC_6117b-768x514.jpg 768w\" sizes=\"(max-width: 800px) 100vw, 800px\" \/><figcaption class=\"wp-element-caption\"><em>Oplismeus burmanni<\/em>.<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"900\" height=\"602\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/03\/DSC_1364.jpg\" alt=\"Elymus canadensis (Canada wild rye)\" class=\"wp-image-454\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/03\/DSC_1364.jpg 900w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/03\/DSC_1364-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/03\/DSC_1364-768x514.jpg 768w\" sizes=\"(max-width: 900px) 100vw, 900px\" \/><figcaption class=\"wp-element-caption\"><em>Elymus canadensis<\/em> (Canada wild rye)<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"900\" height=\"602\" src=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0092.jpg\" alt=\"Hordeum murinum in Copenhagen\" class=\"wp-image-1092\" srcset=\"https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0092.jpg 900w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0092-300x201.jpg 300w, https:\/\/keyapa.com\/sp\/wp-content\/uploads\/2025\/06\/DSC_0092-768x514.jpg 768w\" sizes=\"(max-width: 900px) 100vw, 900px\" \/><figcaption class=\"wp-element-caption\"><em>Hordeum murinum<\/em> in Copenhagen<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\"><strong>References:<\/strong><br><br>Petersen, K. B., and E. A. Kellogg. 2022. Diverse ecological functions and the convergent evolution of grass awns. American Journal of Botany. 109(9): 1331\u20131345. <a href=\"https:\/\/doi.org\/10.1002\/ajb2.16060\">https:\/\/doi.org\/10.1002\/ajb2.16060<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\"><\/p>\n","protected":false},"excerpt":{"rendered":"<p>If one were in the habit of identifying grasses, then the presence of visible and distinctive structures on the spikelets would be one way to ease the pain of such an endeavor. Fortunately, many grasses have long projections from these reproductive structures that go a long way towards helping the observer. These so-called awns are&#8230;<\/p>\n","protected":false},"author":1,"featured_media":3713,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_siteseo_robots_primary_cat":"none","footnotes":""},"categories":[102,9,7],"tags":[141,37,35,71,139,65,23,70,92,81,122,140],"class_list":["post-3701","post","type-post","status-publish","format-standard","has-post-thumbnail","hentry","category-education","category-macrophotography","category-research","tag-aegylops","tag-andropogon","tag-aristida","tag-arrhenatherum","tag-avena","tag-bromus","tag-elymus","tag-hordeum","tag-nassella","tag-oplismenus","tag-piptochaetium","tag-sorghastrum"],"_links":{"self":[{"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/posts\/3701","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/comments?post=3701"}],"version-history":[{"count":25,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/posts\/3701\/revisions"}],"predecessor-version":[{"id":3739,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/posts\/3701\/revisions\/3739"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/media\/3713"}],"wp:attachment":[{"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/media?parent=3701"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/categories?post=3701"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/keyapa.com\/sp\/wp-json\/wp\/v2\/tags?post=3701"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}