{"id":161,"date":"2024-05-10T10:22:36","date_gmt":"2024-05-10T14:22:36","guid":{"rendered":"https:\/\/keyapa.com\/pheidole\/?p=161"},"modified":"2024-05-29T16:57:34","modified_gmt":"2024-05-29T20:57:34","slug":"why-pheidole","status":"publish","type":"post","link":"https:\/\/keyapa.com\/pheidole\/2024\/05\/10\/why-pheidole\/","title":{"rendered":"The Road to Pheidole"},"content":{"rendered":"\n<p>I&#8217;ve always been interested in polymorphic ants, whether it was the mass raiding <em>Carebara diversa<\/em> and its ilk in the Philippines, or the weird top heavy <em>Acanthomyrmex <\/em> species in the tropics. So an attraction to the mostly dimorphic <em>Pheidole<\/em> genus would have been natural.<\/p>\n\n\n\n<br><br>\n\n\n\n<figure data-wp-context=\"{&quot;imageId&quot;:&quot;69f0e4ba87d36&quot;}\" data-wp-interactive=\"core\/image\" data-wp-key=\"69f0e4ba87d36\" class=\"wp-block-image aligncenter size-full wp-lightbox-container\"><img loading=\"lazy\" decoding=\"async\" width=\"750\" height=\"544\" data-wp-class--hide=\"state.isContentHidden\" data-wp-class--show=\"state.isContentVisible\" data-wp-init=\"callbacks.setButtonStyles\" data-wp-on--click=\"actions.showLightbox\" data-wp-on--load=\"callbacks.setButtonStyles\" data-wp-on-window--resize=\"callbacks.setButtonStyles\" src=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/post-2673-0-25573900-1714997927.jpg\" alt=\"\" class=\"wp-image-163\" srcset=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/post-2673-0-25573900-1714997927.jpg 750w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/post-2673-0-25573900-1714997927-300x218.jpg 300w\" sizes=\"auto, (max-width: 750px) 100vw, 750px\" \/><button\n\t\t\tclass=\"lightbox-trigger\"\n\t\t\ttype=\"button\"\n\t\t\taria-haspopup=\"dialog\"\n\t\t\taria-label=\"Enlarge\"\n\t\t\tdata-wp-init=\"callbacks.initTriggerButton\"\n\t\t\tdata-wp-on--click=\"actions.showLightbox\"\n\t\t\tdata-wp-style--right=\"state.imageButtonRight\"\n\t\t\tdata-wp-style--top=\"state.imageButtonTop\"\n\t\t>\n\t\t\t<svg xmlns=\"http:\/\/www.w3.org\/2000\/svg\" width=\"12\" height=\"12\" fill=\"none\" viewBox=\"0 0 12 12\">\n\t\t\t\t<path fill=\"#fff\" d=\"M2 0a2 2 0 0 0-2 2v2h1.5V2a.5.5 0 0 1 .5-.5h2V0H2Zm2 10.5H2a.5.5 0 0 1-.5-.5V8H0v2a2 2 0 0 0 2 2h2v-1.5ZM8 12v-1.5h2a.5.5 0 0 0 .5-.5V8H12v2a2 2 0 0 1-2 2H8Zm2-12a2 2 0 0 1 2 2v2h-1.5V2a.5.5 0 0 0-.5-.5H8V0h2Z\" \/>\n\t\t\t<\/svg>\n\t\t<\/button><figcaption class=\"wp-element-caption\"><em>Carebara diversa<\/em> with Ponerine ant prey in the Philippines, 2004<\/figcaption><\/figure>\n\n\n\n<p>I remember when I was still a teen in the Philippines, and reading in the college library about the rapacious <em>Pheidole megacephala<\/em>, as it devastated native populations of <em>Odontomachus<\/em> in those islands that it had invaded. This was one of the factors that caused me to sally forth to far-off parks to observe interesting ant species. Most times, I took a series of buses and jeepneys for three hours to get to the <a href=\"https:\/\/makiling.center\/the-makiling-botanic-gardens\/\" target=\"_blank\" rel=\"noreferrer noopener\">Mt. Makiling Botanical Garden in Los Banos<\/a>, skipping my college classes in search of more exciting adventures.  It was in these parks that I got to first meet awesome polymorphic ants like <em>Carebara diversa<\/em>, as well as the smaller ants like <em>Pheidole<\/em>.<\/p>\n\n\n\n<p>Fast forward to when I was a student at USC in Los Angeles in the early 1990s. <em>Linepithema humile<\/em> was everywhere, with a good sprinkling of <em>Solenopsis<\/em> fire ants around as well (I don&#8217;t believe it was <em>S. invicta<\/em>, as that did not arrive until the late 1990s). But in the midst of these ants, I found small colonies of a <em>Pheidole<\/em> sp on the USC campus itself, which I treasured and tended to, giving them small bits of food and wishing them well. I always figured it was a precarious existence for these tiny ants, especially when surrounded by such fearsome invaders.<\/p>\n\n\n\n<br><br>\n\n\n\n<figure data-wp-context=\"{&quot;imageId&quot;:&quot;69f0e4ba88295&quot;}\" data-wp-interactive=\"core\/image\" data-wp-key=\"69f0e4ba88295\" class=\"wp-block-image aligncenter size-full is-resized wp-lightbox-container\"><img loading=\"lazy\" decoding=\"async\" width=\"800\" height=\"600\" data-wp-class--hide=\"state.isContentHidden\" data-wp-class--show=\"state.isContentVisible\" data-wp-init=\"callbacks.setButtonStyles\" data-wp-on--click=\"actions.showLightbox\" data-wp-on--load=\"callbacks.setButtonStyles\" data-wp-on-window--resize=\"callbacks.setButtonStyles\" src=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/DSC00632.jpg\" alt=\"\" class=\"wp-image-172\" style=\"width:650px;height:auto\" srcset=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/DSC00632.jpg 800w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/DSC00632-300x225.jpg 300w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/DSC00632-768x576.jpg 768w\" sizes=\"auto, (max-width: 800px) 100vw, 800px\" \/><button\n\t\t\tclass=\"lightbox-trigger\"\n\t\t\ttype=\"button\"\n\t\t\taria-haspopup=\"dialog\"\n\t\t\taria-label=\"Enlarge\"\n\t\t\tdata-wp-init=\"callbacks.initTriggerButton\"\n\t\t\tdata-wp-on--click=\"actions.showLightbox\"\n\t\t\tdata-wp-style--right=\"state.imageButtonRight\"\n\t\t\tdata-wp-style--top=\"state.imageButtonTop\"\n\t\t>\n\t\t\t<svg xmlns=\"http:\/\/www.w3.org\/2000\/svg\" width=\"12\" height=\"12\" fill=\"none\" viewBox=\"0 0 12 12\">\n\t\t\t\t<path fill=\"#fff\" d=\"M2 0a2 2 0 0 0-2 2v2h1.5V2a.5.5 0 0 1 .5-.5h2V0H2Zm2 10.5H2a.5.5 0 0 1-.5-.5V8H0v2a2 2 0 0 0 2 2h2v-1.5ZM8 12v-1.5h2a.5.5 0 0 0 .5-.5V8H12v2a2 2 0 0 1-2 2H8Zm2-12a2 2 0 0 1 2 2v2h-1.5V2a.5.5 0 0 0-.5-.5H8V0h2Z\" \/>\n\t\t\t<\/svg>\n\t\t<\/button><figcaption class=\"wp-element-caption\">Natural habit in the wild of that weird human subspecies, <em>Homo sapiens<\/em> subsp. myrmecofanatico. Here we see the specimen blithely examining an ant nest as tourists pass by and wonder what the heck he&#8217;s doing.<\/figcaption><\/figure>\n\n\n\n<p>In college, I also frequented the library, and one of the journals I read was Psyche: A Journal of Entomology. In this wonderful journal I came upon two of the coolest ant species ever. Both were <em>Pheidole<\/em>, and reading about them stoked my interest in the genus like never before. The first species was <em>Phedole ridicula<\/em> (now <em>P. absurda<\/em>), and the article was <a href=\"https:\/\/www.hindawi.com\/journals\/psyche\/1966\/081641\/\" target=\"_blank\" rel=\"noreferrer noopener\"><em>The Habits of Pheidole Ridicula Wheeler, With Remarks on Habit Patterns in the Genus Pheidole<\/em><\/a>. This study dealt mainly with the amazing major of this species and its efficacy in defending the nest. The second article was also authored by Creighton, and called <a href=\"https:\/\/www.hindawi.com\/journals\/psyche\/1959\/020849\/\" target=\"_blank\" rel=\"noreferrer noopener\"><em>The Habits of Pheidole Militicida Wheeler<\/em><\/a>. It also elaborated on the defensive function of the major of the equally fascinating <em>Pheidole militicida<\/em>.<\/p>\n\n\n\n<br><br>\n\n\n\n<figure data-wp-context=\"{&quot;imageId&quot;:&quot;69f0e4ba8873e&quot;}\" data-wp-interactive=\"core\/image\" data-wp-key=\"69f0e4ba8873e\" class=\"wp-block-image size-full wp-lightbox-container\"><img loading=\"lazy\" decoding=\"async\" width=\"800\" height=\"472\" data-wp-class--hide=\"state.isContentHidden\" data-wp-class--show=\"state.isContentVisible\" data-wp-init=\"callbacks.setButtonStyles\" data-wp-on--click=\"actions.showLightbox\" data-wp-on--load=\"callbacks.setButtonStyles\" data-wp-on-window--resize=\"callbacks.setButtonStyles\" src=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/Pheidole_militicida_casent0102884_profile_1.jpg\" alt=\"\" class=\"wp-image-115\" srcset=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/Pheidole_militicida_casent0102884_profile_1.jpg 800w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/Pheidole_militicida_casent0102884_profile_1-300x177.jpg 300w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/Pheidole_militicida_casent0102884_profile_1-768x453.jpg 768w\" sizes=\"auto, (max-width: 800px) 100vw, 800px\" \/><button\n\t\t\tclass=\"lightbox-trigger\"\n\t\t\ttype=\"button\"\n\t\t\taria-haspopup=\"dialog\"\n\t\t\taria-label=\"Enlarge\"\n\t\t\tdata-wp-init=\"callbacks.initTriggerButton\"\n\t\t\tdata-wp-on--click=\"actions.showLightbox\"\n\t\t\tdata-wp-style--right=\"state.imageButtonRight\"\n\t\t\tdata-wp-style--top=\"state.imageButtonTop\"\n\t\t>\n\t\t\t<svg xmlns=\"http:\/\/www.w3.org\/2000\/svg\" width=\"12\" height=\"12\" fill=\"none\" viewBox=\"0 0 12 12\">\n\t\t\t\t<path fill=\"#fff\" d=\"M2 0a2 2 0 0 0-2 2v2h1.5V2a.5.5 0 0 1 .5-.5h2V0H2Zm2 10.5H2a.5.5 0 0 1-.5-.5V8H0v2a2 2 0 0 0 2 2h2v-1.5ZM8 12v-1.5h2a.5.5 0 0 0 .5-.5V8H12v2a2 2 0 0 1-2 2H8Zm2-12a2 2 0 0 1 2 2v2h-1.5V2a.5.5 0 0 0-.5-.5H8V0h2Z\" \/>\n\t\t\t<\/svg>\n\t\t<\/button><figcaption class=\"wp-element-caption\"><em>Pheidole militicida<\/em>&nbsp;from Antweb. Copyright Jen Fogarty<\/figcaption><\/figure>\n\n\n\n<p>But notwithstanding my interest in these Pheidole species, my main love for the next few years were for the<a href=\"https:\/\/keyapa.com\/leafcutters\/\" target=\"_blank\" rel=\"noreferrer noopener\"> <em>Atta<\/em> leafcutter ants<\/a>. These species are strongly polymorphic, with incredibly large and fearsome major workers, and I even took trips to places like Texas specifically to search for them. I had a particularly strong liking for species that had huge shiny-headed soldiers, like the <em>Atta mexicana<\/em> that I found in Mexico.<\/p>\n\n\n\n<br><br>\n\n\n\n<figure data-wp-context=\"{&quot;imageId&quot;:&quot;69f0e4ba88c25&quot;}\" data-wp-interactive=\"core\/image\" data-wp-key=\"69f0e4ba88c25\" class=\"wp-block-image aligncenter size-full is-resized wp-lightbox-container\"><img loading=\"lazy\" decoding=\"async\" width=\"600\" height=\"422\" data-wp-class--hide=\"state.isContentHidden\" data-wp-class--show=\"state.isContentVisible\" data-wp-init=\"callbacks.setButtonStyles\" data-wp-on--click=\"actions.showLightbox\" data-wp-on--load=\"callbacks.setButtonStyles\" data-wp-on-window--resize=\"callbacks.setButtonStyles\" src=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/asj_atta_cephalotes1.jpg\" alt=\"\" class=\"wp-image-183\" style=\"width:650px;height:auto\" srcset=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/asj_atta_cephalotes1.jpg 600w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/asj_atta_cephalotes1-300x211.jpg 300w\" sizes=\"auto, (max-width: 600px) 100vw, 600px\" \/><button\n\t\t\tclass=\"lightbox-trigger\"\n\t\t\ttype=\"button\"\n\t\t\taria-haspopup=\"dialog\"\n\t\t\taria-label=\"Enlarge\"\n\t\t\tdata-wp-init=\"callbacks.initTriggerButton\"\n\t\t\tdata-wp-on--click=\"actions.showLightbox\"\n\t\t\tdata-wp-style--right=\"state.imageButtonRight\"\n\t\t\tdata-wp-style--top=\"state.imageButtonTop\"\n\t\t>\n\t\t\t<svg xmlns=\"http:\/\/www.w3.org\/2000\/svg\" width=\"12\" height=\"12\" fill=\"none\" viewBox=\"0 0 12 12\">\n\t\t\t\t<path fill=\"#fff\" d=\"M2 0a2 2 0 0 0-2 2v2h1.5V2a.5.5 0 0 1 .5-.5h2V0H2Zm2 10.5H2a.5.5 0 0 1-.5-.5V8H0v2a2 2 0 0 0 2 2h2v-1.5ZM8 12v-1.5h2a.5.5 0 0 0 .5-.5V8H12v2a2 2 0 0 1-2 2H8Zm2-12a2 2 0 0 1 2 2v2h-1.5V2a.5.5 0 0 0-.5-.5H8V0h2Z\" \/>\n\t\t\t<\/svg>\n\t\t<\/button><figcaption class=\"wp-element-caption\"><em>Atta cephalotes<\/em> soldier<\/figcaption><\/figure>\n\n\n\n<p>By 2017 I had started a <a href=\"https:\/\/pheidolemegacephala.blogspot.com\/\" target=\"_blank\" rel=\"noreferrer noopener\">site dedicated to the invasive <em>Pheidole megacephala<\/em><\/a>, and during trips to tropical regions I would invariably spend some hours surveying the area for this species. While my wife and our friends went off on shore excursions, I would be busy trudging along the touristy streets and byways, my eyes glued to the ground and scanning ceaselessly for the 2 mm long minor workers.<\/p>\n\n\n\n<br><br>\n\n\n\n<figure data-wp-context=\"{&quot;imageId&quot;:&quot;69f0e4ba890fc&quot;}\" data-wp-interactive=\"core\/image\" data-wp-key=\"69f0e4ba890fc\" class=\"wp-block-image size-full wp-lightbox-container\"><img loading=\"lazy\" decoding=\"async\" width=\"771\" height=\"511\" data-wp-class--hide=\"state.isContentHidden\" data-wp-class--show=\"state.isContentVisible\" data-wp-init=\"callbacks.setButtonStyles\" data-wp-on--click=\"actions.showLightbox\" data-wp-on--load=\"callbacks.setButtonStyles\" data-wp-on-window--resize=\"callbacks.setButtonStyles\" src=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/keywest.png\" alt=\"\" class=\"wp-image-166\" srcset=\"https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/keywest.png 771w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/keywest-300x199.png 300w, https:\/\/keyapa.com\/pheidole\/wp-content\/uploads\/2024\/05\/keywest-768x509.png 768w\" sizes=\"auto, (max-width: 771px) 100vw, 771px\" \/><button\n\t\t\tclass=\"lightbox-trigger\"\n\t\t\ttype=\"button\"\n\t\t\taria-haspopup=\"dialog\"\n\t\t\taria-label=\"Enlarge\"\n\t\t\tdata-wp-init=\"callbacks.initTriggerButton\"\n\t\t\tdata-wp-on--click=\"actions.showLightbox\"\n\t\t\tdata-wp-style--right=\"state.imageButtonRight\"\n\t\t\tdata-wp-style--top=\"state.imageButtonTop\"\n\t\t>\n\t\t\t<svg xmlns=\"http:\/\/www.w3.org\/2000\/svg\" width=\"12\" height=\"12\" fill=\"none\" viewBox=\"0 0 12 12\">\n\t\t\t\t<path fill=\"#fff\" d=\"M2 0a2 2 0 0 0-2 2v2h1.5V2a.5.5 0 0 1 .5-.5h2V0H2Zm2 10.5H2a.5.5 0 0 1-.5-.5V8H0v2a2 2 0 0 0 2 2h2v-1.5ZM8 12v-1.5h2a.5.5 0 0 0 .5-.5V8H12v2a2 2 0 0 1-2 2H8Zm2-12a2 2 0 0 1 2 2v2h-1.5V2a.5.5 0 0 0-.5-.5H8V0h2Z\" \/>\n\t\t\t<\/svg>\n\t\t<\/button><figcaption class=\"wp-element-caption\">Map of <em>Pheidole megacephala<\/em> in Key West, FL (Sunjian, 2018)<\/figcaption><\/figure>\n\n\n\n<p>In a way, it was almost natural that I ultimately expanded the scope of my interest to the entire genus <em>Pheidole<\/em>, after my obsession with <em>P. megacephala<\/em>. Now I am not limited to studying just one species, albeit a species that is so fascinating, but learning about the evolution and lifestyles of the most diverse ant genus in the world. It&#8217;ll be a fun adventure.<\/p>\n\n\n\n<p><strong>Literature Cited<\/strong><\/p>\n\n\n\n<p>Sunjian, A (2018).<a href=\"https:\/\/pheidolemegacephala.blogspot.com\/2018\/06\/a-survey-of-ants-in-old-town-key-west.html\" target=\"_blank\" rel=\"noreferrer noopener\"> A survey of ants in Old Town Key West in Florida and a review of Pheidole megacephala as an archetypal \u201chegemonic\u201d species. The Pm Compendium (PMC).<\/a> <\/p>\n\n\n\n<p><\/p>\n\n\n\n<p><\/p>\n\n\n\n<p><\/p>\n\n\n\n<p><\/p>\n\n\n\n<p><\/p>\n","protected":false},"excerpt":{"rendered":"<p>I&#8217;ve always been interested in polymorphic ants, whether it was the mass raiding Carebara diversa and its ilk in the Philippines, or the weird top heavy Acanthomyrmex species in the tropics. So an attraction to the mostly dimorphic Pheidole genus would have been natural. I remember when I was still a teen in the Philippines,&#8230;<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_siteseo_robots_primary_cat":"","footnotes":""},"categories":[10],"tags":[],"class_list":["post-161","post","type-post","status-publish","format-standard","hentry","category-personal-notes"],"_links":{"self":[{"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/posts\/161","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/comments?post=161"}],"version-history":[{"count":29,"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/posts\/161\/revisions"}],"predecessor-version":[{"id":294,"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/posts\/161\/revisions\/294"}],"wp:attachment":[{"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/media?parent=161"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/categories?post=161"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/keyapa.com\/pheidole\/wp-json\/wp\/v2\/tags?post=161"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}